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20 reviewsSUMMARYCells are essential to understanding health and disease, yet traditional models fall short of modeling andsimulating their function and behavior. Advances in AI and omics offer groundbreaking opportunities tocreate an AI virtual cell (AIVC), a multi-scale, multi-modal large-neural-network-based model that can represent and simulate the behavior of molecules, cells, and tissues across diverse states. This Perspective provides a vision on their design and how collaborative efforts to build AIVCs will transform biological researchby allowing high-fidelity simulations, accelerating discoveries, and guiding experimental studies, offeringnew opportunities for understanding cellular functions and fostering interdisciplinary collaborations inopen science.INTRODUCTION